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44. Family DOLICHOPODIDAE

[Originally] by D.J. Bickel & C.E. Dyte

Flies of the family Dolichopodidae are often abundant in moist habitats and are frequently found on foliage, tree trunks, mud flats, and river rocks. Adults are predaceous on soft-bodied invertebrates and serve as important general control agents of many pest species.

The maggotlike larvae are found in such habitats as soil, rotted vegetation, mud, and under bark. Most are predators or scavengers, although the larva of Thrypticus is a phytophagous stem miner in various Monocotyledons [see Dyte (1967) for review of immature stages]. Williams' (1938b,c, 1939) studies of the immature stages of Hawaiian dolichopodids are among the best documented for the family.

The elaborate secondary sexual characters of male dolichopodids are assumed to aid in species recognition during courtship and often show parallel development in unrelated groups. These characters include flaglike flattening of the arista and tarsi, strongly modified setae and cuticular projections, prolongation and deformation of podomeres, orientated silvery pruinosity, maculation and deformation of wings, etc. The hypopygium is often greatly enlarged and exposed and usually carries species-specific characters. The tiny Babindella, of the endemic Australian subfamily Babindellinae, shows secondary postabdominal symmetry (Bickel 1987b).

The Dolichopodidae is one of the most diverse families of flies, with approximately 5,000 described species. In the Australasian/Oceanian Regions, 995 species representing 77 genera in 9 subfamilies are currently known, although many new species and genera await description, especially from Australia and the Melanesian Archipelago. Apart from southern temperate elements in New Zealand and Australia, much of the Australasian/Oceanian fauna has strong affinities with that of the Oriental Region. A notable exception is the presence of the Neotropical Condylostylus longicornis in eastern Polynesia.

Some "tramp" species, such as Medetera grisescens and Chrysosoma leucopogon, show extraordinarily widespread distributions throughout both the Oriental and Australasian regions, including many isolated oceanic islands. Several brachypterous species are known from subantarctic islands. Of particular interest is the extensive speciation of Campsicnemus in the Hawaiian Islands (138 spp., with many more awaiting description). This radiation invites comparison with that of the well-studied Hawaiian Drosophila.

The subfamily arrangements followed here are based on Ulrich (1981), with modifications. The limits of certain subfamilies, such as the Diaphorinae and Sympycninae are unclear and some genera, such as Sympycnus, are polyphyletic assemblages. Recent revisionary studies on the Medeterinae (Bickel 1986a,b,c, 1987c) and Hydrophorinae (Grootaert & Meuffels 1984, Meuffels & Grootaert 1984) have added to the regional understanding of these taxa.

Becker's (1922b) monograph, although dated, remains the basic reference for both the Oriental and Australasian/Oceanian regions. Important faunistic treatments include those on the Hawaiian Islands (Hardy & Kohn 1964, Tenorio 1969), New Zealand (Parent 1933b), Australia (Hardy 1930a, 1935b, 1939c; Parent 1932a, 1933a), Samoan Islands (Lamb 1929), and New Guinea (de Meijere 1906b, 1913a,b, 1915b; Parent 1939a). Additional species descriptions are found in the many papers of Parent. Robinson & Vockeroth (1981) provided a well-illustrated general review of the family.

Subfamily SCIAPODINAE

Genus ABBEMYIA Bickel

ABBEMYIA Bickel, 1994: 212. Type species: Psilopus nigrofsciatus Macquart, 1850, orig. des.
nigrofasciata Macquart, 1850: 430(126) (Psilopus). Australia, "côte orientale"; Australia (NSW, Qld).

regale Parent, 1932a: 111 (Chrysosoma). Australia (NSW).

chetiscutatum Parent, 1932a: 113 (Chrysosoma). Australia (NSW).
taree Bickel, 1994: 214. Australia (NSW).

Genus AMBLYPSILOPUS Bigot

AMBLYPSILOPUS Bigot, 1888d: xxiv. Type species: Psilopus psittacinus Loew, 1861 (as psitacinus Fabricius), orig. des.
GNAMPTOPSILOPUS Aldrich, 1893: 48. Type species: Psilopus scintillans Loew, 1861, des. Coquillett, 1910: 547.
AUSTRALIOLA Parent, 1932a: 127. Type species: Australiola tonnoiri Parent, 1932 [= Sciapus zonatus Parent, 1932], mon.
AGONOSOMA, authors, not Guérin-Méneville, 1838, misid.
CHRYSOSOMA, authors, not Guérin-Méneville, 1838, misid.
CONDYLOSTYLUS, authors, not Bigot, 1859, misid.
PSILOPUS, authors, not Meigen, 1824, misid.
SCIAPUS, authors, not Zeller, 1842, misid.
abruptus Walker, 1859c: 115 (Psilopus). Indonesia (Sulawesi); Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG); Christmas I (Ocean I), Cocos (Keeling) Is, India, Indonesia (Java, Sumatra), Malaysia (Pen, Sarawak), Philippines, Singapore, Taiwan, Vietnam.

muticus Thomson, 1869: 509 (Psilopus). Cocos (Keeling) Is.

filatus Wulp, 1884d: 227 (Psilopus). Indonesia (Java).

recurrens de Meijere, 1913b: 342 (Psilopus). Indonesia (Irian Jaya).
albicinctus de Meijere, 1913a: 56 (Psilopus). Indonesia (Maluku).
albipes Becker, 1922b: 201 (Sciapus). PNG (PNG).
albisignatus Bickel, 1994: 294. Australia (NSW); Australia (Qld).
aliciensis Bickel, 1994: 357. Australia (NT).
alter Becker, 1922b: 202 (Sciapus). PNG (PNG).
annanensis Bickel, 1994: 291. Australia (Qld).
anomalicornis Becker, 1922b: 199 (Sciapus). Australia (NT); Australia (Qld, WA).
arboreus Bickel, 1994: 304. Australia (NT).
argyrodendron Bickel, 1994: 320. Australia (Qld).
augustus Bickel, 1994: 331. Australia (Qld).
austerus Parent, 1935a: 363 (Chrysosoma). Malaysia (Sabah); Guam, Northern Marianas.
babindensis Bickel, 1994: 349. Australia (Qld).
baroalba Bickel, 1994: 329. Australia (NT).
basistylatus Parent, 1939a: 164 (Sciapus). PNG (PNG).
basseti Bickel, 1994: 302. Australia (Qld); Australia (NSW).
bataviensis Bickel, 1994: 351. Australia (Qld).
belauensis Bickel, 1995: 104. Belau; Micronesia.
berensis Bickel, 1994: 331. Australia (NSW): Australia (Qld).
bertiensis Bickel, 1994: 333. Australia (Qld).
bimus Bickel, 1994: 298. Australia (Qld).
biprovincialis Bickel, 1994: 309. Australia (NT); Australia (Qld), Micronesia.
birraduk Bickel, 1994: 329. Australia (NT).
borroloola Bickel, 1994: 344. Australia (NT); Australia (Qld).
brevitibia Bickel, 1994: 304. Australia (NSW).
byrnei Bickel, 1994: 294. Australia (Qld).
cahillensis Bickel, 1994: 341. Australia (NT).
callainus Bickel, 1994: 336. Australia (NT).
canungra Bickel, 1994: 297. Australia (Qld); Australia (NSW).
careelensis Bickel, 1994: 319. Australia (NSW); Australia (Qld), Norfolk I.
cincinnatus Bickel, 1994: 357. Australia (Qld).
cobourgensis Bickel, 1994: 347. Australia (NT).
commoni Bickel, 1994: 283. Australia (WA); Australia (NT, Qld).
cooki Bickel, 1994: 329. Australia (Qld); Australia (NT).
cosmochirus Bezzi, 1928: 61 (Chrysosoma). Fiji; Tonga.
cursus Bickel, 1994: 315. Australia (NSW).
cyplus Bickel, 1994: 350. Australia (Qld).
dohni Bickel, 1994: 355. Australia (Qld).
edwardsi Bickel, 1994: 339. Australia (NT).
eupulvillatus Parent, 1928: 194 (Condylostylus). Tonga.
flaviappendiculatus de Meijere, 1910a: 94 (Agonosoma). Indonesia (Java); Australia (NT, Qld); China, Indonesia (Nusa Tenggara), Philippines, Vietnam.

dolichonemis Frey, 1925: 17 (Psilopus). Philippines.

brevitarsis Parent, 1932b: 110 (Sciapus). Indonesia (Nusa Tenggara).
eotrogon Bickel, 1994: 336. Australia (Qld).
fonsecai Bickel, 1994: 341. Australia (NSW); Australia (Qld).
fonticola Bickel, 1994: 286 (as fonticolus). Australia (NT).
fortescuia Bickel, 1994: 284. Australia (WA).
fustis Bickel, 1994: 321. Australia (Qld).
gapensis Bickel, 1994: 351. Australia (Qld).
glaciunguis Bickel, 1994: 323. Australia (Qld).
graciliventris Parent, 1933a: 178 (Sciapus). Australia (Qld); Australia (NT, WA).
greenwoodi Bezzi, 1928: 61 (Chrysosoma). Fiji.
gressitti Bickel, 1994: 311. Australia (NT).
guntheri Bickel, 1994: 317. Lord Howe I; Australia (NSW).
humilis Becker, 1922b: 172 (Chrysosoma). Taiwan; Solomon Is, Western Samoa; China, India, Malaysia (Pen), Nepal, Philippines.

sauteri Becker, 1924c: 127 (Chrysosoma). Taiwan.
julius Bickel, 1994: 309. Australia (Qld).
jullatensis Bickel, 1994: 323. Australia (Qld).
kakaduensis Bickel, 1994: 323. Australia (NT).
kaputar Bickel, 1994: 303. Australia (NSW).
liepae Bickel, 1994: 317. Lord Howe I.
lismorensis Bickel, 1994: 299. Australia (NSW).
loriensis Bickel, 1994: 299. Australia (NSW).
malensis Bickel, 1994: 283. Australia (Qld); Australia (NSW).
melasma Bickel, 1994: 359. Australia (NT).
mensualis Bickel, 1994: 343. Australia (Qld); Australia (NT).
moggillensis Bickel, 1994: 302. Australia (Qld).
mollis Parent, 1932a: 125 (Sciapus). Australia (NSW); Australia (Qld).
montanorum Bickel, 1994: 306. Australia (NSW); Australia (ACT).
nambourensis Bickel, 1994: 278. Australia (Qld); Australia (NSW).
neoplatypus Bickel, 1994: 277 (n.n. for platypus Parent). Indonesia (Maluku); Australia (Qld), PNG (PNG).

platypus Parent, 1932e: 350 [1932f: 580] (Sciapus). Indonesia (Maluku). [Preocc. Becker, 1922.]
nimbuwah Bickel, 1994: 284. Australia (NT).
pallidicornis Grimshaw, 1901: 12 (Gnamptopsilopus). Hawaiian Is; Belau, French Polynesia (Society Is), Guam, PNG (PNG); Seychelles, Taiwan.

fulgidipenne Enderlein, 1912c: 377 (Chrysosoma). Taiwan.
papilliferus Bickel, 1994: 280. Australia (Qld); Australia (NT).
parallelinervis Parent, 1935c: 75 (Sciapus). Solomon Is.
paramonovi Bickel, 1994: 317. Lord Howe I.
pectoralis de Meijere, 1913a: 54 (Psilopus). Indonesia (Maluku).
ponapensis Bickel, 1995: 102. Micronesia.
pseudexsul Bickel, 1994: 312. Australia (NSW).
pulverulentus Parent, 1939a: 162 (Chrysosoma). PNG (PNG).
pulvillatus Bezzi, 1928: 68 (Condylostylus). Fiji.
putealis Bickel, 1994: 317. Australia (WA).
quldensis Bickel, 1994: 343. Australia (Qld).
rentzi Bickel, 1994: 303. Australia (NSW).
rimbija Bickel, 1994: 346. Australia (NT); Australia (Qld).
sabroskyi Bickel, 1995: 105. Belau.
septentrionalis Bickel, 1994: 280. Australia (Qld).
sideroros Bickel, 1994: 359. Australia (Qld); PNG (PNG).
superans Walker, 1860b: 150 (Psilopus). Indonesia (Maluku); Indonesia (Irian Jaya), PNG (PNG).

fluminis de Meijere, 1913b: 344 (Psilopus). Indonesia (Irian Jaya).
tenuitarsis de Meijere, 1913a: 343 (Psilopus). Indonesia (Irian Jaya).
tinarooensis Bickel, 1994: 334. Australia (Qld).
topendensis Bickel, 1994: 344. Australia (NT).
tortus Bickel, 1994: 306. Australia (Qld).
tozerensis Bickel, 1994: 282. Australia (Qld).
triduum Bickel, 1994: 320. Australia (Qld).
triscuticatus Hardy, 1930a: 129 (Sciapus). Australia (Qld); Australia (ACT, NSW).
trogon Bickel, 1994: 335. Australia (WA).
trudis Bickel, 1994: 297. Australia (Qld).
trukensis Bickel, 1995: 107. Micronesia.
uneorum Bickel, 1994: 296. Australia (Qld).
uptoni Bickel, 1994: 328. Australia (NT).
variabilis de Meijere, 1913a: 346 (Psilopus). Indonesia (Irian Jaya); PNG (PNG).
walkeri Bickel, 1994: 296. Australia (Qld).
webbensis Bickel, 1994: 328. Australia (Qld).
wellsae Bickel, 1994: 348. Australia (NT).
williamsi Bickel, 1994: 311. Australia (NSW); Australia (Qld).
wokoensis Bickel, 1994: 300. Australia (NSW).
wongabelensis Bickel, 1994: 348. Australia (Qld).
zonatus Parent, 1932a: 123 (Sciapus). Australia (NSW); Australia (Qld), Lord Howe I.

tonnoiri Parent, 1932a: 128 (Australiola). Australia (NSW).

Genus AUSTROSCIAPUS Bickel

AUSTROSCIAPUS Bickel, 1992: 13 (key) [1994: 231 (full description)]. Type species: Sciapus proximus Parent, 1928, mon. [by orig. des. in Bickel 1994].
actensis Bickel, 1994: 241. Australia (ACT).
aprilis Bickel, 1994: 254. Australia (Qld).
ascitus Bickel, 1994: 246. Australia (Qld).
balli Bickel, 1994: 267. Australia (Qld).
bifarius Becker, 1922b: 208 (Sciapus). Australia (Qld) [not India]; Australia (NT).
broulensis Bickel, 1994: 252. Australia (NSW); Auatralia (Qld).
cantrelli Bickel, 1994: 262. Australia (Qld); Australia (NSW).
capricornis Bickel, 1994: 244. Australia (Qld).
cassisi Bickel, 1994: 245. Australia (NSW).
collessi Bickel, 1994: 239. Australia (Qld).
connexus Walker, 1835a: 471 (Psilopus). "New Holland"; Australia (NSW, Qld, WA), French Polynesia (Austral Is, Society Is), Hawaiian Is, Norfolk I.

pachigyna Macquart, 1848: 197(37) (Psilopus). Australia (NSW).

pachygyna, error for pachigyna.
crater Bickel, 1994: 248. Australia (Qld).
dayi Bickel, 1994: 261. Australia (NSW); Australia (Qld).
dekeyzeri Bickel, 1994: 265. Australia (NSW).
dendrohalma Bickel, 1994: 261. Australia (Qld); Australia (NSW).
discretifasciatus Macquart, 1850: 431(127) (Psilopus). Australia (Tas); Australia (ACT, NSW, Qld, Vic).

depinctus Becker, 1922b: 210. Australia (NSW).

genevieveae Parent, 1932a: 168 (as genevievei). Australia (Qld).
doddi Bickel, 1994: 239. Australia (Qld).
flavicauda Bickel, 1994: 257. Australia (NSW).
frauci Bickel, 1994: 241. Australia (Qld).
fraudulosus Bickel, 1994: 239. Australia (Qld).
gwynnae Bickel, 1994: 257. Australia (Qld); Australia (NSW).
hollowayi Bickel, 1994: 268. Australia (WA).
janae Bickel, 1994: 263. Australia (NSW); Australia (Qld, Vic).
magus Bickel, 1994: 246. Australia (NSW); Australia (Qld).
minnamurra Bickel, 1994: 248. Australia (NSW).
muelleri Bickel, 1994: 258. Australia (NSW).
nellae Bickel, 1994: 264. Australia (NSW).
otfordensis Bickel, 1994: 259. Australia (NSW).
proximus Parent, 1928: 191 (Sciapus). Australia (NSW); Australia (Qld), NZ (NZ).
pseudotumidus Bickel, 1994: 252. Australia (Qld).
pulvillus Bickel, 1994: 270. Australia (WA).
quadrimaculatus Parent, 1932a: 121 (Sciapus). Australia (NSW).
ravenshoensis Bickel, 1994: 249. Australia (Qld).
riparius Bickel, 1994: 247. Australia (NSW).
sarinensis Bickel, 1994: 255. Australia (Qld).
solus Bickel, 1994: 267. Australia (NSW).
stevensi Bickel, 1994: 256. Australia (NSW).
storeyi Bickel, 1994: 265. Australia (Qld).
tooloomensis Bickel, 1994: 257. Australia (NSW): Australia (Qld).
triangulifer Becker, 1922b: 211 (Sciapus). Australia (Qld) [not India].

insecans Becker, 1922b: 208 (Sciapus). Australia (Qld) [not India].
tumidus Hardy, 1959: 299 (Sciapus). Australia (NSW); Australia (ACT, Qld, Vic).
zentae Bickel, 1994: 241. Australia (NSW); Australia (Qld).

Genus CHRYSOSOMA Guérin-Méneville

CHRYSOSOMA Guérin-Méneville, 1831: pl. 20, fig. 6. Type species: Chrysosoma fasciata Guérin-Méneville, 1831, des. Enderlein, 1912c: 373.
AGONOSOMA Guérin-Méneville, 1838: 293 (unnec. n. n. for Chrysosoma). [Preocc. Laporte, 1832.]
MARGARITOSTYLUS Bigot, 1859a: 215, 222. Type species: Psilopus globifer Wiedemann, 1830, orig. des.
MEGISTOSTYLUS Bigot, 1859a: 215, 222. Type species: Dolichopus crinicornis Wiedemann, 1824 (as Psilopus), orig. des.
OARIOSTYLUS Bigot, 1859a: 215, 223. Type species: Psilopus tuberculicornis Macquart, 1855, orig. des.
MEGYSTISTYLUS. Incorr. orig. spell. of Megistostylus (Bigot, 1859a: 222).
OARIOPHERUS Bigot, 1888d: xxiv. Type species: Psilopus tuberculicornis Macquart, 1855, orig. des.
SPATHIOPSILOPUS Bigot, 1888d: xxiv. Type species: Psilopus globifer Wiedemann, 1830, orig. des.
SPATHIPSILOPUS Bigot, 1890b: 268. Type species: Psilopus globifer Wiedemann, 1830, des. Dyte, 1975: 213.
CONDYLOSTYLUS, authors, not Bigot, 1859, misid.
GNAMPTOPSILOPUS, authors, not Aldrich, 1893, misid.
PSILOPUS, authors, not Meigen, 1824, misid.
PSILOPODINUS, authors, not Bigot, 1888, misid.
MEGISTOTYLUS error for Megistostylus.
aeneum Fabricius, 1805: 268 (Dolichopus). Indonesia (Java); Indonesia (Irian Jaya, Maluku), PNG (PNG).

variipennis Walker, 1861b: 238 (Psilopus). Indonesia (Irian Jaya).
agrihasn Bickel, 1995: 87. Northen Marianas.
aldrichi de Meijere, 1913b: 342 (Psilopus). Indonesia (Irian Jaya); PNG (PNG).

collini Parent, 1932g: 217. PNG (PNG).
antennatum Becker, 1922b: 149. PNG (Bismarck Arch).

trichromatum Parent, 1939a: 163. PNG (Bismarck Arch).
arrogans Parent, 1934b: 113. Solomon Is; Guam, PNG (Bougainville I); Philippines.
austeni Parent, 1934a: 14. PNG (PNG); Indonesia (Irian Jaya). [Proposed (incorrectly) as a n. n. for specimens misidentified as lilacinus var. flavitibiale de Meijere. Name made available by back reference to Becker, 1922b: 182.]
bicolor Parent, 1937a: 133. Solomon Is.
callosum Parent, 1929: 199. Australia (Qld); Australia (NT).
ceramense de Meijere, 1913a: 55. Indonesia (Maluku).
clarkei Bickel, 1995: 88. Micronesia.
complicatum Becker, 1922b: 159. Marshall Is; American Samoa, Fiji, Kiribati (Gilbert Is), Marshall Is, Micronesia, Nauru, Palmyra Atoll, Tonga, Tuvalu, Wake I, Western Samoa.

leucochirum Bezzi, 1928: 64. Fiji.
compressipes Parent, 1939a: 157. PNG (PNG).
confusum Parent, 1932g: 226. PNG (PNG. [Proposed (incorrectly) as a n. n. for specimens misidentified as Chrysosoma papuasinum Bigot. Name made available by back reference to Becker, 1922b: 179.]
consimile Lamb, 1929: 133. Western Samoa; American Samoa.
cordieri Parent, 1934c: 279. Indonesia (Java); PNG (Bismarck Arch).
crinicorne Wiedemann, 1824: 39 (Dolichopus). Indonesia (Java); Australia (Qld), Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, PNG), Solomon Is; ?Brazil, Ryukyu Is, widesp. Oriental tropics.

filifer Walker, 1859b: 114 (Psilopus). Indonesia (Sulawesi).

longisetosus Wulp, 1882: 120 (Psilopus). Brazil.

aeterus Bigot, 1890b: 283 (Psilopus). Indonesia (Maluku).

imparile Parent, 1933a: 174. Australia (Qld).

longicornis, authors, not Fabricius, 1775, misid.
crypticum Becker, 1922b: 193. PNG (Bismarck Arch).
digitulum Becker, 1922b: 190. PNG (PNG).
dilectum Parent, 1935a: 368. Malaysia (Sabah); PNG (PNG).
disparitarse Parent, 1935c: 66. Solomon Is.

nigricinctum Parent, 1941: 203. Solomon Is.
diversicolor Parent, 1928: 197. Australia (Qld).
duplociliatum Parent, 1933a: 172. Australia (NT); Australia (Qld).
egens Walker, 1858b: 92 (Psilopus). Indonesia (Maluku); Indonesia (Irian Jaya).
eminens Parent, 1935c: 68. Solomon Is; PNG (PNG).
excellens Parent, 1934b: 114. Solomon Is.
fasciatum Guérin-Méneville, 1831: pl. 20 [1838: 293]. Indonesia (Irian Jaya); Indonesia (Maluku).

cognatum Parent, 1929: 200. Indonesia (Maluku).
ferriferum Lamb, 1929: 126. Western Samoa; American Samoa, Fiji, Tonga.

liber Parent, 1934c: 293 (Condylostylus). Fiji.
fissilamellatum Parent, 1939a: 158. PNG (PNG).
flavitibiale de Meijere, 1915b: 119 (Psilopus lilacinus ssp.). Indonesia (Irian Jaya).
formosum Parent, 1934c: 281. Not given [?Australasian].
gaumense Bickel, 1995: 81. Guam.
globiferum Wiedemann, 1830a: 222 (Psilopus). China; Hawaiian Is; Hong Kong, Ryukyu Is, Taiwan.

figuratum Becker, 1922b: 184. Hong Kong.

fraternum Van Duzee, 1933: 310. Hawaiian Is.
ignavum Becker, 1922b: 174. PNG (PNG).
impressum Becker, 1922b: 173. PNG (PNG); Indonesia (Irian Jaya).
inerme de Meijere, 1913b: 343 (Psilopus albopilosus ssp.). Indonesia (Irian Jaya); Australia (Qld), PNG (PNG).
inflatum Becker, 1922b: 221 (Condylostylus). PNG (PNG).
innatum Lamb, 1929: 134. Western Samoa.
interruptum Becker, 1922b: 177. Australia (NT).
kusaiense Bickel, 1995: 85. Micronesia.
lacteimicans Becker, 1923: 28. "Samoa"; Western Samoa.

decorum Lamb, 1929: 132. Western Samoa.

latefuscatum Parent, 1929: 204. "Iles Samoa."
latemaculatum Parent, 1939a: 160. PNG (PNG).
leopoldi Parent, 1932h: 23. Indonesia (Irian Jaya).
leucopogon Wiedemann, 1824: 40 (Dolichopus). "India orient"; Australia (NT, Qld), Belau, French Polynesia (Society Is), New Caledonia (New Caledonia), PNG (PNG), Western Samoa; Chistmas I (Indian Ocean), Cocos (Keeling) Is, Madagascar, Mascarenes, Seychelles, Taiwan, Tanzania, widesp. Oriental tropics (incl. Chagos Is).

apicalis Wiedemann, 1830a: 227 (Psilopus). Indonesia (Sumatra).

conicornis Macquart, 1846: 248(120) (Psilopus). India.

curviseta Thomson, 1869: 508 (Psilopus). French Polynesia (Society Is).

loewi Enderlein, 1912c: 378. Taiwan.
leucopygum de Meijere, 1906b: 82 (Agonosoma). Indonesia (Irian Jaya); PNG (PNG).
leveri Parent, 1934b: 115. Solomon Is.

festivum Parent, 1935c: 69. Solomon Is.
lilacinum de Meijere, 1913b: 341 (Psilopus). Indonesia (Irian Jaya).
lucare Bickel, 1994: 204. Australia (Qld); Australia (NT, WA).
lucigena Walker, 1858b: 91 (Psilopus). Indonesia (Maluku); Indonesia (Irian Jaya), PNG (PNG).

extendens Walker, 1865b: 111 (Psilopus). "New Guinea" [= Indonesia (Irian Jaya)].
luctuosum Parent, 1928: 195. Fiji.
ludens Parent, 1935c: 70. Solomon Is; Marshall Is, PNG (Bismarck Arch, PNG).
maculipenne Guérin-Méneville, 1831: pl. 20 [1838: 294]. Indonesia (?Sumatra); Indonesia (Irian Jaya, Maluku), PNG (PNG).

benedictus Walker, 1858b: 91 (Psilopus). Indonesia (Maluku).

persuadens Walker, 1860b: 149 (Psilopus). Indonesia (Maluku).

diversifrons de Meijere, 1913b: 345 (Psilopus). Indonesia (Irian Jaya).
maculiventre Parent, 1935c: 71. Solomon Is.

leuocpygum, Parent, 1934, not de Meijere, 1906, misid.
marginale Walker, 1861b: 283 (Psilopus). Indonesia (Maluku); Indonesia (Irian Jaya), PNG (PNG).

obscuripennis de Meijere, 1913a: 54 (Psilopus). Indonesia (Maluku).

anthracinum Becker, 1922b: 191. PNG (PNG).
mariana Bickel, 1995: 87. Northern Marianas.
medium Becker, 1922b: 139. PNG (PNG).
meijeri Parent, 1932e: 349 [1932f: 579]. Indonesia (Maluku); Malaysia (Sabah).
molestum Parent, 1934c: 288. Vanuatu; Kiribati (Gilbert Is), Tuvalu.
mutilatum Parent, 1935c: 72. Solomon Is; Micronesia.
nigrohalteratum Parent, 1939a: 161. PNG (PNG).
nigrolimbatum de Meijere, 1913b: 340 (Psilopus). Indonesia (Irian Jaya).
nobile Parent, 1933a: 175. Australia (NT); Australia (Qld, WA).
noumeanum Bigot, 1890b: 288 (Psilopus). New Caledonia (New Caledonia).

bigoti Parent, 1932g: 223. New Caledonia (New Caledonia).
obscuripes Parent, 1934b: 117. Solomon Is; PNG (Bougainville I).
pacificum Parent, 1930: 113. French Polynesia (Society Is).

globiferum, authors, not Wiedemann, 1830, misid.
pagdeni Parent, 1937a: 138. Solomon Is.
palapes Hardy & Kohn, 1964: 234. Hawaiian Is; Sri Lanka.
papuasinum Bigot, 1890b: 283 (Spathiospilopus). "Nouvelle-Guinée"; Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG).
parvicucullatum Lamb, 1929: 129. Western Samoa.
patelliferum Thomson, 1869: 507 (Psilopus). Guam.

macropus Thomson, 1869: 508 (Psilopus). Guam.
pelagica Bickel, 1995: 81. Philippines; Guam.
pexum Becker, 1922b: 173. PNG (PNG).

pexoides Parent, 1939a: 161. PNG (PNG).
planitarse Becker, 1922b: 168. PNG (PNG).
proliciens Walker, 1856c: 120 (Psilopus). Malaysia (Sarawak); Australia (NT, Qld), Indonesia (Irian Jaya, Maluku), PNG (PNG); India, Indonesia, Philippines, Singapore.

delectans Walker, 1856c: 120 (Psilopus). Malaysia (Sarawak).

perficiens Walker, 1860b: 150 (Psilopus). Indonesia (Maluku).

albopilosus Wulp, 1895a: 46 (Psilopus). Indonesia (Java).

divisum Becker, 1922b: 182. PNG (PNG) & India & Malaysia & Philippines.

diversum. Incorr. orig. spell. of divisum (Becker, 1922b: 242).

altopilosum, error for albopilosus.
provocans Parent, 1934c: 289. Vanuatu.
pseudocallosum Bickel, 1994: 193. Australia (NT); Australia (WA), PNG (PNG).
purpurascens de Meijere, 1915b: 117 (Psilopus; n. n. for purpuratum de Meijere). Indonesia (Irian Jaya); PNG (PNG).

purpuratum de Meijere, 1906b: 84 (Agonosoma). Indonesia (Irian Jaya). [Preocc. Aldrich, 1901.]
quadratum Wulp, 1884d: 222 (Psilopus). Indonesia (Maluku).
salomone Parent, 1929: 205. Solomon Is.
saonekense de Meijere, 1913a: 56 (Psilopus). Indonesia (Maluku); PNG (PNG).

atripes Parent, 1934c: 278. Indonesia (Maluku).
seticorne Walker, 1864: 234 (Psilopus). Indonesia (Maluku).
sigma Parent, 1935c: 73. Solomon Is.
solitarium Parent, 1935c: 74. Solomon Is.
splendidum Wulp, 1869: 111 (Psilopus). Indonesia (Irian Jaya); PNG (PNG).

signatipenne de Meijere, 1906b: 83 (Agonosoma). Indonesia (Irian Jaya).
townesi Bickel, 1995: 84. Micronesia.
trigemmans Walker, 1849: 650 (Psilopus). Not given [?Australasian].
tuberculicorne Macquart, 1855: 104(84) (Psilopus). French Polynesia (Marquesas); French Polynesia (Society Is), Niue.

candidum Parent, 1932d: 872. French Polynesia (Society Is).
viduum Lamb, 1929: 135. Western Samoa.
waigeense de Meijere, 1913a: 57 (Psilopus). Indonesia (Maluku); Indonesia (Irian Jaya), PNG (PNG).

geniculatum de Meijere, 1913b: 347 (Psilopus). Indonesia (Irian Jaya).

appendiculatus de Meijere, 1915b: 120 (Psilopus geniculatus ssp.). Indonesia (Irian Jaya). [Preocc. Bigot, 1890.]

mutabile Becker, 1922b: 173 (n. n. for appendiculatus de Meijere).

futile Parent, 1939a: 158. PNG (PNG).
yapense Bickel, 1995: 83. Micronesia.

Genus CONDYLOSTYLUS Bigot

CONDYLOSTYLUS Bigot, 1859a: 215, 223. Type species: Psilopus bituberculatus Macquart, 1842, orig. des.
TYLOCHAETUS Bigot, 1888d: xxiv. Type species: Psilopus bituberculatus Macquart, 1842, orig. des.
AGONOSOMA, authors, not Guérin-Méneville, 1838, misid.
CHRYSOSOMA, authors, not Guérin-Méneville, 1838, misid.
PSILOPUS, authors, not Meigen, 1824, misid.
SCIAPUS, authors, not Zeller, 1842, misid.
longicornis Fabricius, 1775: 783 (Musca). "America"; French Polynesia (Austral Is, Marquesas Is, Society Is, Tuamotu Arch); Neotrop. Reg., USA.

chrysoprasi Walker, 1849: 646 (Psilopus). "West Indies."

iliipes Aldrich, 1901: 355 (Psilopus). Mexico.

Genus DYTOMYIA Bickel

DYTOMYIA Bickel, 1994: 91. Type species: Sciapus sordidus Parent, 1928, orig. des.
bancrofti Bickel, 1994: 96. Australia (Qld).
flaviseta Bickel, 1994: 98. Australia (WA).
sordidus Parent, 1928: 193 (Sciapus). Australia (NSW); Australia (ACT, Qld, SA, Vic).

anomalipennis Hardy, 1930: 128 (Sciapus). Australia (Qld).
torresiana Bickel, 1994: 96. Australia (Qld).
tumifrons Bickel, 1994: 97. Australia (WA); Australia (NT).

Genus HELIXOCERUS Lamb

HELIXOCERUS Lamb, 1929: 137. Type species: Helixocerus mendosum Lamb, 1929, orig. des.
mendosum Lamb, 1929: 137. American Samoa; Western Samoa.

Genus HETEROPSILOPUS Bigot

HETEROPSILOPUS Bigot, 1859a: 215, 224. Type species: Psilopus grandis Macquart, 1850 [= Heteropsilopus cingulipes Walker, 1835], orig. des.
CHRYSOSOMA, authors, not Guérin-Méneville, 1838, misid.
PSILOPUS, authors, not Meigen, 1824, misid.
SCIAPUS, authors, not Zeller, 1842, misid.
araluensis Bickel, 1994: 174. Australia (NSW); Australia (ACT).
brevicornis Macquart, 1850: 428(124) (Psilopus). Australia (Tas); Australia (NSW, Qld, SA, Vic, WA).

venustus Walker, 1852: 209 (Psilopus). Australia (WA).

volucre Becker, 1922b: 142 (Chrysosoma). "Neu Holland" & "Australien."

bimaculatus Parent, 1932a: 117 (Sciapus). Australia (NSW).
brindabellensis Bickel, 1994: 176. Australia (ACT); Australia (NSW, Vic).
caelicus Parent, 1932a: 167 (Chrysosoma). Australia (Qld).
calabyi Bickel, 1994: 178. Australia (Vic).
cingulipes Walker, 1835a: 471 (Psilopus). "New Holland"; widesp. Australia (excl. NT, WA).

sydneyensis Macquart, 1846: pl. 11, fig. 16 [1847a: 39; 1847b: 55 (as sidneyensis)] (Psilopus ). Australia (NSW).

grandis Macquart, 1850: 430(126) (Psilopus). Australia (NSW).

eximius Walker, 1852: 209 (Psilopus). "New Holland."

chrysurgus Schiner, 1868: 214 (Psilopus). Australia (NSW).

angulosus Bigot, 1890b: 285 (Psilopus). "Australie."

chalceus White, 1916b: 250 (Sciapus). Australia (Tas).

alatum Becker, 1922b: 188 (Chrysosoma). Australia (NSW).

micans Parent, 1932a: 109 (Chrysosoma). Australia (ACT).

metallicum Parent, 1932a: 113 (Chrysosoma). Australia (NSW).

jacquelineae Parent, 1932a: 169 (Sciapus; as jacquelinei). Australia (ACT).

alatrum, error for alatum.
ingenuus Erichson, 1842: 273 (Psilopus). Australia (Tas); Australia (ACT, NSW, Qld, SA, Vic).

gloriosus Parent, 1932a: 119. Australia (Tas).
intermedius Bickel, 1994: 179. Australia (SA).
khooi Bickel, 1994: 167. Australia (NSW); Australia (Qld).
meensis Bickel, 1994: 171. Australia (Qld).
plumifer Becker, 1922b: 206 (Sciapus). Australia (NSW); Australia (Qld).
savicensis Bickel, 1994: 175. Australia (SA).
squamifer Hardy, 1958: 295. Australia (NSW).
sugdeni Bickel, 1994: 165. Australia (NSW).
tantanoola Bickel, 1994: 176. Australia (SA).
trifasciatus Macquart, 1850: 430(126) (Psilopus). Australia (Tas); Australia (NSW, SA, Vic, WA).
tweedensis Bickel, 1994: 170. Australia (Qld); Australia (NSW).

Genus KRAKATAUIA Enderlein

KRAKATAUIA Enderlein, 1912c: 408. Type species: Psilopus rectus Wiedemann, 1830 (as Pilopus), orig. des.
CHRYSOSOMA, authors, not Guérin-Méneville, 1838, misid.
CONDYLOSTYLUS, authors, not Bigot, 1859, misid.
SCIAPUS, authors, not Zeller, 1842, misid.
abbreviata Becker, 1922b: 193 (Chrysosoma). PNG (Bismarck Arch).
alanae Bickel, 1994: 152. Australia (NSW); Australia (Qld).
barbescens Parent, 1939a: 156 (Chrysosoma). PNG (PNG).
claudiensis Bickel, 1994: 151. Australia (Qld); PNG (PNG).
evulgata Becker, 1922b: 205 (Sciopus). PNG (Bismarck Arch); Marshall Is, Micronesia, Nauru, PNG (PNG), Solomon Is, Vanuatu.

frontale Becker, 1922b: 185 (Chrysosoma aldrichi ssp.). PNG (PNG).

pressipes Parent, 1929: 244 (Sciapus). Micronesia [not Nearctic].

occultus Parent, 1934b: 124 (Sciapus). Solomon Is.
funeralis Parent, 1933a: 173 (Chrysosoma). Australia (Qld); Australia (NSW).
macalpinei Bickel, 1994: 146. Australia (NSW); Australia (Qld).
malanda Bickel, 1994: 153. Australia (Qld).
melanochira Bezzi, 1928: 63 (Chrysosoma). Fiji.
micronesiana Bickel, 1995: 99. Guam; Belau, Midway Atoll, Northern Marianas, Wake I.
nitidifacies Parent, 1934b: 123 (Sciapus). PNG (Bismarck Arch).
nupta Bezzi, 1928: 67 (Condylostylus). Fiji.
obversicornis Bickel, 1994: 146. Australia (Qld).
pseudofuneralis Bickel, 1994: 151. Australia (Qld).
recta Wiedemann, 1830a: 225 (Psilopus). Indonesia (Sumatra); PNG (PNG); Oriental Reg.

unitus Parent, 1928: 193 (Sciapus). Malaysia (Sarawak).
remota Bickel, 1994: 155. Australia (Qld).
sericea de Meijere, 1913a: 57 (Psilopus). Indonesia (Maluku).
tabulina Becker, 1922b: 197 (Sciapus). PNG (PNG).
trustorum Bickel, 1994: 155. Australia (NSW).

Genus MESORHAGA Schiner

MESORHAGA Sc